A new study published in Cell on January 5 reconstructs the genetic history of Scandinavia over the past 2,000 years, from the Iron Age to the present.This look back at the history of Scandinavia is based on an analysis of 48 new and 249 previously published ancient human genomes representing numerous iconic archaeological sites, as well as genetic data from more than 16,500 living Scandinavians today.

The new study conducted by Stockholm University and deCODE genetics (Reykjavik) provides insight into migration patterns and gene flux during the Viking era, among other intriguing findings (750–1050 CE).

It also indicates that ancestries introduced to the region during the Viking era declined over time for unknown reasons.

Ricardo Rodrguez-Varela of Stockholm University stated, “Although non-local ancestry is still evident in modern Scandinavians, levels of non-local ancestry in some regions are lower than those observed in ancient individuals from the Viking to Medieval periods.”

This suggests that ancient individuals with non-Scandinavian ancestry contributed proportionally less to the present gene pool in Scandinavia than would have been predicted based on patterns observed in the archaeological record.

“Multiple mechanisms introduced individuals from various regions to Scandinavia [at various periods],” added Anders Gotherstrom of Stockholm University.

The researchers had not initially intended to link together the history of Scandinavia across time and space. Instead, they were engaged in three distinct investigations focusing on distinct archaeological sites.

“When analyzing the genetic affinities of individuals from different archaeological sites, such as the Vendel period boat burials, Viking period chamber burials, and well-known archaeological sites like the Migration period Sandby borg ringfort, known for the mssacre that occurred there [in] 500 CE, as well as individuals from the 17th-century royal Swedish warship Kronan, we began to observe differences in the levels and origins of non-local ancestry across the different archaeological sites.”

Archaeological excavation at Sandby Borg.

“Initially, we were working with three different studies,” Götherström said. “One on Sandby borg, one on the boat burials, and one on the man-of-war Kronan. At some point it made more sense to unite them to one study on the Scandinavian demography during the latest 2,000 years.”

The goal was to document how past migrations have affected the Scandinavian gene pool across time and space to better understand the current Scandinavian genetic structure.

As reported in the new study, the researchers found regional variation in the timing and magnitude of gene flow from three sources: the eastern Baltic, the British Irish Isles, and southern Europe.

British Irish ancestry was widespread in Scandinavia from the Viking period, whereas eastern Baltic ancestry is more localized to Gotland and central Sweden. In some regions, a drop in current levels of external ancestry suggests that ancient immigrants contributed proportionately less to the modern Scandinavian gene pool than indicated by the ancestry of genomes from the Viking and Medieval periods.

Finally, the data show that a north-south genetic cline that characterizes modern Scandinavians is mainly due to differential levels of Uralic ancestry. It also shows that this cline existed in the Viking Age and possibly even earlier.

Götherström suggests that what the data reveal about the nature of the Viking period is perhaps most intriguing. The migration from the west impacted all of Scandinavia, and the migration from the east was Sєx biased, with movement primarily of female people into the region.

As the researchers write, the findings overall “indicate a major increase [in gene flow] during the Viking period and a potential bias toward females in the introduction of eastern Baltic and, to a lesser extent, British-Irish ancestries.

Underwater excavations of the ship Kronan.

“Gene flow from the British-Irish Isles during this period seems to have had a lasting impact on the gene pool in most parts of Scandinavia,” they continued.

“This is perhaps not surprising given the extent of Norse activities in the British-Irish Isles, starting in the 8th century with recurrent raids and culminating in the 11th century North Sea Empire, the personal union that united the kingdoms of Denmark, Norway, and England. The circumstances and fate of people of British-Irish ancestry who arrived in Scandinavia at this time are likely to have been variable, ranging from the forced migration of slaves to the voluntary immigration of more high-ranking individuals such as Christian missionaries and monks.”

Overall, the findings show that the Viking period in Scandinavia was a very dynamic time, they say, with people moving around and doing many different things. In future work, they hope to add additional genetic data in hopes of learning more about how the ancestries that arrived during the Viking period were later diluted.

They’d also like to pinpoint when the north-south cline was shaped based on study of larger ancient datasets from the north.

“We need more pre-Viking individuals form north Scandinavia to investigate when the Uralic ancestry enter in this region,” Rodríguez-Varela said.

“Also, individuals from 1000 BCE to 0 are very scarce, [and] retrieving DNA from Scandinavian individuals with these chronologies will be important to understand the transition from the Bronze Age to the Iron Age in this part of the world. Finally, more individuals from the Medieval period until the present will help us to understand when and why we observe a reduction in the levels of non-local ancestry in some current regions of Scandinavia.”

“There is so much fascinating information about our prehistory to be explored in ancient genomes,” Götherström said.

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